Origin of Species & Continent FormationA The fact that there was once a Pangean supercontinent, a Panthalassa
Ocean, and a Tethys Ocean, has profound implications for the evolution
of multicellular life on Earth. These considerations were unknown to the
scientists of the 19th century—making their scientific deductions even more
remarkable. Quite independently of each other, Charles Darwin and his
young contemporary Alfred Russel Wallace reached the conclusion that life
had evolved by natural selection. Wallace later wrote in My Life of his own
inspiration:B Why do some species die and some live? The answer was clearly that on
the whole the best fitted lived. From the effects of disease the most healthy
escaped;from enemies the strongest, the swiftest or the most cunning from
famine the best hunters…then it suddenly flashed on me that this self-acting
process would improve the race, because in every generation the inferior
would inevitably be killed off and the superior would remain, that is , the
fittest
would survive.C Both Darwin’s and Wallace’s ideas about natural selection had been influenced
by the essays of Thomas Malthus in his Principles of Population. Their
conclusions, however, had been the direct result of their personal observation
of animals and plants in widely separated geographic locations:Darwin from
his experiences during the voyage of the Beagle, and particularly during the
ship’s visit to the Galapagos Islands in the East Pacific in 1835; Wallace during
his years of travel in the Amazon Basin and
in the Indonesia-Australian Archipelago in
the 1850s.
D Darwin had been documenting his ideas on
natural selection for many years when he
received a paper on this selfsame subject
60
EF
from Wallace, who asked for Darwin’s opinion and help in getting it published.
In July 1858, Charles Lyell and J. D Hooker, close friends of Darwin, pressed
Darwin to present his conclusions so that he would not lose priority to
and unknown naturalist. Presiding over the hastily called but now historic
meeting of the Linnean Society in London, Lyell and Hooker explained to the
distinguished members how
“these two gentlemen” (who were absent:Wallace
was abroad and Darwin chose not to attend), had
“independently and unknown
to one another, conceived the same very ingenious theory,
”B oth Darwin and Wallace had realized that the anomalous distribution
of species in particular regions had profound evolutionary significance.
Subsequently, Darwin spent the rest of his days in almost total seclusion
thinking and writing mainly about the origin of species. In contrast, Wallace
applied himself to the science of biogeography, the study of the pattern and
distribution of species, and its significance, resulting in the publication of a
massive two-volume work the Geographical Distribution of Animals in 1876.W allace was a gentle and modest man, but also persistent and quietly
courageous. He spent years working in the most arduous possible climates and
terrains, particularly in the Malay archipelago, he made patient and detailed
zoological observations and collected huge number of specimens for museums
and collectors-which is how he made a living. One result of his work was
the conclusion that there is a distinct faunal boundary, called
“Wallace’s line,
“between an Asian realm of animals in Java, Borneo and the Philipiones and
an Australian realm in New Guinea and Australia. In essence this boundary
posed a difficult question: How on Earth did plants and animals with a clear
affinity to the Northern Hemisphere meet with their Southern Hemispheric
counterparts along such a distinct Malaysian demarcation zone? Wallace was
uncertain about demarcation on one particular island-Celebes, a curiously
shaped place that is midway between the two groups. Initially he assigned its
flora-fauna to the Australian side of the line, but later he transferred it to the
61
Asian side. Today we know the reason for his dilemma. 200MYA East and
West Celebes were islands with their own natural history lying on opposite
sides of the Tethys Ocean. They did not collide until about 15 MYA. The
answer to the main question is that Wallace’s Line categorizes Laurasia-derived flora-fauna (the Asian) and Gondwana-derived flora-fauna (the
Australian), fauna that had evolved on opposing shares of the Tethys. The
closure of the Tethys Ocean today is manifested by the ongoing collision of
Australia/New Guinea with Indochina/Indonesia and the continuing closure of
the Mediterranean Sea—a remnant of the Western Tethys Ocean.
G In his origin of continents and oceans, Wegener quoted at length from
Wallace’s Geographical Distribution of Animals. According to Wegener’s
reading, Wallace had identified three clear divisions of Australian animals,
which supported his own theory of continental displacement. Wallace had
shown that animals long established in southwestern Australia had an affinity
with animals in South Africa, Madagascar, India, and Ceylon, but did not have
an affinity with those in Asia. Wallace also showed that Australian marsupials
62
HI
and monotremes are clearly related to
those in South America, the Moluccas,
and various Pacific islands, and that none
are found in neighboring Indonesia. From
this and related data, Wegener concluded that the then broadly accepted
“landbridge” theory could not account for this distribution of animals and that
only his theory of continental drift could explain it.
T he theory that Wegener dismissed in preference to his own proposed that
plants and animals had once migrated across now-submerged intercontinental
landbridges. In 1885, one of Europe’ s leading geologists, Eduard Suess,
theorized that as the rigid Earth cools, its upper crust shrinks and wrinkles like
the withering skin of an aging apple. He suggested that the planet’s seas and
oceans now fill the wrinkles between once-contiguous plateaus.T oday, we know that we live on a dynamic Earth with shifting, colliding and
separating tectonic plates, not a
“withering skin”, and the main debate in the
field of biogeography has shifted. The discussion now concerns
“ dispersalism”
versus
“vicarianism: unrestricted radiation of species on the one hand and the
development of barriers to migration on the other. Dispersion is a short-term
phenomenon—the daily or seasonal migration of species and their radiation
to the limits of their natural environment on an extensive and continuous
landmass. Vicarian evolution, however, depends upon the separation and
isolation of a variety of species within the confines of natural barriers in the
form of islands, lakes, or shallow seas—topographical features that take a long
time to develop.63
Questions 1-5
.................................................................................Use the information in the passage to match the people (listed A-E) with opinions or
deeds below. Write the appropriate letters A-E in boxes 1-5 on your answer you may use any letter more than onceA
B
C
D
E
1
2
3
4
5
SuessWallaceDarwin and WallaceWegenerLyell and HookerPersuade Darwin to publish his scientific findingsDepicted physical feature of earth’s uced continental drift hed works about wildlife distribution in different ion of species is based on selection by ons 6-8
.................................................................................The reading Passage has nine paragraphs paragraph contains the following information?Write the correct letter A-I, in boxes 6-8 on your answer sheet.6
7
8
Best adaptable animal survived on the ry called Wallace’s line found between Asia and relevance exists between Australia and Africa.64
Questions 9-13
...............................................................................SummaryComplete the following summary of the paragraphs of Reading Passage, using no more
than words from the Reading Passage for each answer. Write your answers in boxes 9-13
on your answer r found that continental drift instead of “land bridge” theory could explain
strange species’ distribution phenomenon. In his theory, vegetation and wildlife
9to 10mobile 11 intercontinentally. However, Eduard Suess compared the wrinkle of crust
of an old apple. Now it is well known that we are living on a constant
instead of what Suess described. Hot spot in biogeography are
”and “13”switched to concerns between two terms: “ 1265
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